its best if they are back in the original bores as they sort of "wear in" togeather, the bore and lifter.
But like Brett said, they will work ok in any hole, unless one is damaged. When you lube them up, try each one and make sure none are to tight or too sloppy.
Use a fair amount of assembly lube on the lifters/bores, the rocker arms and the cam bearings. Don't be shy. And don't just use engine oil, the assembly lube is readily availiable and cheap. A new head ruined due to damaged cam caps is not.
The cam caps on the other hand MUST be back in the correct location and orientation. They are marked however, so triple check. Your head and possibly the cams too, will be damaged if you run it with the cam caps on incorrectly. The torque is also very small, 12-14lbs/ft